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The Nuclear Structures of Protocaryotic Organisms (Bacteria by G. Wolfgang Fuhs (auth.)

By G. Wolfgang Fuhs (auth.)

of energids in line with telephone is believed to equivalent the variety of genetic enhances in addition to the variety of conceivable cells that finally could emerge from it with no replication of its genetic fabric. (In eucaryotic cells, polyenergidy happens within the type of cells containing a number of nuclei every one or as polyploidy, concerning the co-existence of genollles in the barriers of a unmarried nucleus. evidently phrases corresponding to "poly nucleated" and "polyploid" arc beside the point for protocaryotic cells.) The variety of energids pCI' telephone will be topic to version as a reaction to yes environmental stipulations 01' in the course of sure levels of a developmental cycle. The absence in protocaryotic nuclear our bodies of structural parts except DNA markedly impacts their constitution and morphology. because the protocaryon primarily is an accumulation of DNA, the quantity, mo­ lecular association and chemical nation of the DNA are easy determinants of nuclear form and fantastic constitution. for this reason, the equipped DNA molecule (the genophor) needs to be thought of the imperative topic of any treatise facing nuclear cytology in micro organism and Cyanophyceae.

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For details see KRISCH 1965, JOSET et al. ) From the fact that establishment of cell-to-cell contact almost immediately results in transfer of the genetic material, JACOB et al. (196'3) speculated that the F replicator system lIlay be located at the cell periphery and in immediate neighborhood of the antigen which presllmably is active in establishing contact with the recipient cell. At this point, considerations on the possible organization of an E. coli Hfr nucleoid during conjugation reflect on models of the nor m a I mode of replication in such cells.

10 according to which, upon D;\"A replication, two nucleoids are fonned in adjacent positions containing one double-stranded unit each. The lIucleoids are distributed into daughter cells according to their intracellular arrangement. A different segregation pattern was observed in succinate-grown cell" of E. RK and BIRD 1965 a). Such cells con tain two genetic complements each which arc replicated alternating and in sequence (K. G. LAHK and C: LAnK 1965). At cell division, each genetic complement contributes half of its progeny to each of the daughter cells.

Coli). and others. Fig. 11. \, Illodel for the regulation of DNA synthcsis and for an equal distribution of DNA among the daughter lmct

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